In 1956 I presented a discussion of the Coleoptera indigenous to the seashore of Pacific North America. I divided the fauna into zones and subzones based on type of shore and reach of tide. Among the genera listed were two of the family Tenebrionidae, Epantius (now called Apsena) and Phaleria. Each of these is represented in southern California by a single species in decaying seaweed on sandy beaches. The first of these,Apsena obscura LeConte, is found only in the dry or nearly dry seaweed left on the berm of the beach by the highest tide of each tidal period (Moore's Zone 2, subzone A). A. obscura is sometimes common but not nearly as common as the second species, Phaleria rotundata LeConte. P. rotundata is unusual among the Coleoptera of this fauna in that it is one of the few species to be commonly encountered in two separate subzones. It is often abundant in both decaying seaweed and dry seaweed (Moore's subzones 2A and 2B). It is one of the most common insects of the southern California seashore.
"Notes on Phaleria Rotundata Leconte with Description of the Larva (Coleoptera: Tenebrionidae),"
The Great Lakes Entomologist, vol 7
Available at: https://scholar.valpo.edu/tgle/vol7/iss4/1