A survey of scarab beetles (Coleoptera: Scarabaeidae) was conducted during 2009 in the geographically distinct area of Winneshiek and Allamakee counties in extreme Northeast Iowa. Four plant communities distributed among 10 locations, including tallgrass prairie, mixed grassland and forest, oak-hickory forest, and pasture were sampled during 2009 using black light traps, banana- beer traps, rodent burrow pitfalls, manure-baited pitfalls, and gleaning. Additionally, previous specimen records were examined. Forty-nine scarab species were documented for Winneshiek and Allamakee counties, including five species not previously reported in Iowa. Mixed grassland and forest habitat contained the greatest scarab species richness (n=31); black light trapping yielded the greatest number of individual beetles and collected the most species.
Worthington, Reese J. and Larsen, Kirk J.
"An Annotated Checklist of Scarab Beetles (Coleoptera: Scarabaeidae) from Northeastern Iowa,"
The Great Lakes Entomologist, vol 43
Available at: https://scholar.valpo.edu/tgle/vol43/iss1/7