A sharply delineated, bilateral differentiation of color patterns and morphology were observed in a final (5th) instar larva of a subspecies backcross of a female Papilio glaucus glaucus with a hybrid male (P. g. glaucus x P. g. canadensis). Color and morphological differences were detectable in the pupa as well. In addition, a bilateral size difference was evident in both the pupa and the resulting adult butterfly. Such observations within a single living individual attest to the bilateral independence (also evident in perfect gynandromorphs) and general flexibility of the developmental control in this species of Lepidoptera.
Scriber, J. Mark and Hagen, Robert H.
"Bilateral Differentiation of Color and Morphology in the Larval and Pupal Stages of Papilio Glaucus (Lepidoptera: Papilionidae),"
The Great Lakes Entomologist, vol 23
Available at: https://scholar.valpo.edu/tgle/vol23/iss3/3