Rearing immatures of Euschistus tristigmus tristigmus in 8L:16D, 8L: 4D:8L;4D, and 16L:8D showed that the length of each photophase, rather than an accumulated total of shorter photophases, during each 24 hr period was the determining factor in producing adult dimorphism in shoulder shape and number of midventral abdominal spots.
McPherson, J. E.
"Effects of Continuous and Split Developmental Photophases During Each 24 Hour Period on Adult Morphology in Euschistus Tristigmus Tristigmus (Hemiptera: Pentatomidae),"
The Great Lakes Entomologist, vol 12
Available at: https://scholar.valpo.edu/tgle/vol12/iss2/4